Dr. Quilliam received his PhD in Biochemistry from the University of Sheffield, England where he studied cell signaling events that regulate pituitary hormone secretion. His postdoctoral training focused on the discovery and characterization of Ras family small GTP binding proteins, their regulators, effectors, and the biological/pathological processes that they control. This work, performed in the laboratories of Drs. Joan Heller Brown (UCSD), Gary Bokoch (The Scripps Research Institute) and Channing Der (La Jolla Cancer Research Foundation and University of North Carolina, Chapel Hill), provided a strong foundation in biochemical and molecular techniques and a disciplined approach to solving biological questions. Dr. Quilliam’s laboratory at IU School of Medicine has continued this tradition since 1995. Using molecular, cellular and in vivo models, his students and postdoctoral fellows have contributed to the discovery and characterization of multiple cell signaling events associated with cell growth and cancer.